This research was supported under an Australian Research Council (ARC) Discovery Project Funding Scheme (DP160101380). M.L. is supported by an ARC Future Fellowship (FT210100656). P.D. is supported by an Australian National Health and Medical Research Council (NHMRC) Senior Research Fellowship (1136908). D.L. is supported by an NHMRC Leadership Fellowship (1196892). J.H. is supported by the Wellcome Trust (grant number: 208090/Z/17/Z). A.P. is supported by an Australian Research Council Discovery Early Career Researcher Award (DE190101074).